{"id":148,"date":"2020-03-31T18:37:25","date_gmt":"2020-03-31T09:37:25","guid":{"rendered":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/?p=148"},"modified":"2021-04-09T19:09:59","modified_gmt":"2021-04-09T10:09:59","slug":"topicrubisco","status":"publish","type":"post","link":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/research\/topicrubisco.html","title":{"rendered":"\u5149\u5408\u6210\u306e\u9375\u9175\u7d20RubisCO\u3092\u7528\u3044\u305f\u3082\u306e\u4f5c\u308a"},"content":{"rendered":"\n<h3>\u5149\u5408\u6210\u3068\u306f<\/h3>\n\n\n\n<p>\u5149\u5408\u6210\u306f\u3001\u5149\u306e\u30a8\u30cd\u30eb\u30ae\u30fc\u3092\u5229\u7528\u3057\u3066\u6c34\u3068\u4e8c\u9178\u5316\u70ad\u7d20\u304b\u3089\u6709\u6a5f\u7269\u3092\u5408\u6210\u3057\u307e\u3059\u3002\u8a00\u3046\u307e\u3067\u3082\u306a\u304f\u3001\u5730\u7403\u4e0a\u306e\u751f\u547d\u6d3b\u52d5\u3092\u7dad\u6301\u3059\u308b\u3046\u3048\u3067\u6700\u3082\u91cd\u8981\u306a\u53cd\u5fdc\u3067\u3059\u3002\u690d\u7269\u3067\u975e\u5e38\u306b\u767a\u9054\u3057\u3066\u3044\u307e\u3059\u304c\u3001\u7d30\u83cc\u306b\u3082\u5149\u5408\u6210\u80fd\u529b\u3092\u6301\u3064\u3082\u306e\u304c\u3044\u307e\u3059\u3002\u5149\u5408\u6210\u306f\u3001\u3069\u306e\u3088\u3046\u306a\u4ed5\u7d44\u307f\u3067\u4e8c\u9178\u5316\u70ad\u7d20\u3092\u6709\u6a5f\u7269\u3078\u5909\u63db\u3059\u308b\u306e\u3067\u3057\u3087\u3046\uff1f\u306a\u3093\u3068\u306a\u304f\u3001\u4e8c\u9178\u5316\u70ad\u7d20\u5206\u5b50\u306b\u5149\u304c\u5f53\u305f\u3063\u3066\u30a8\u30cd\u30eb\u30ae\u30fc\u304c\u4e0e\u3048\u3089\u308c\u308b\u306e\u304b\u306a\u3068\u601d\u3063\u3066\u3044\u307e\u305b\u3093\u304b\uff1f\u5b9f\u306f\u3001\u5149\u5408\u6210\u3067\u306f\u3001\u5149\u30a8\u30cd\u30eb\u30ae\u30fc\u3092\u53d7\u5bb9\u3057\u3066\u5316\u5b66\u30a8\u30cd\u30eb\u30ae\u30fc\u306b\u5909\u63db\u3059\u308b\u5149\u5316\u5b66\u7cfb\u3068\u3001\u4e8c\u9178\u5316\u70ad\u7d20\u3092\u6709\u6a5f\u7269\u306b\u5909\u63db\u3059\u308b\u30ab\u30eb\u30d3\u30f3\u30b5\u30a4\u30af\u30eb\u3068\u547c\u3070\u308c\u308b\u7d4c\u8def\u306f\u5225\u306e\u30b7\u30b9\u30c6\u30e0\u3067\u3059\u3002\u3067\u3059\u306e\u3067\u3001\u4e00\u90e8\u306e\u5fae\u751f\u7269\u306f\u3001\u5149\u304c\u7121\u304f\u3066\u3082\u70ad\u9178\u56fa\u5b9a\u304c\u3067\u304d\u307e\u3059\u3002\u4f7f\u7528\u3059\u308b\u9084\u5143\u529b\u306f\u3001\u6c34\u7d20\u3084\u9244\u306a\u3069\u69d8\u3005\u3067\u3059\u3002<br>\u3000\u4e8c\u9178\u5316\u70ad\u7d20\u304c\u6709\u6a5f\u7269\u306b\u5909\u63db\u3055\u308c\u308b\u53cd\u5fdc\u3092\u89e6\u5a92\u3059\u308b\u306e\u306f\u3001 Ribulose 1,5-bisphosphate (RuBP) carboxylase\/oxygenase (RubisCO)\u3068\u3044\u3046\u9175\u7d20\u3067\u3059\u3002\u3053\u306e\u9175\u7d20\u306fRuBP(\u70ad\u7d20\u6570\uff15)\u3068CO2\u304b\u30893-phosphoglycerate\uff08\u70ad\u7d20\u6570\uff13\uff09\u30922\u5206\u5b50\u751f\u6210\u3057\u307e\u3059\u3002\u3064\u307e\u308a\u3001\u70ad\u7d20\u6570\u306b\u7740\u76ee\u3059\u308b\u3068\u30015+1=3&#215;2\u3068\u3044\u3046\u53cd\u5fdc\u3067\u3059\u3002\u3053\u306e\u53cd\u5fdc\u306f\u3001ATP\u306a\u3069\u306e\u30a8\u30cd\u30eb\u30ae\u30fc\u4f9b\u7d66\u306a\u3057\u306b\u9032\u884c\u3057\u307e\u3059\u3002CO2\u306e\u56fa\u5b9a\u5316\u53cd\u5fdc\u305d\u306e\u3082\u306e\u306f\u30a8\u30cd\u30eb\u30ae\u30fc\u304c\u3044\u3089\u306a\u3044\u306e\u3067\u3059\u3002\u306a\u305c\u305d\u3093\u306a\u3053\u3068\u304c\u53ef\u80fd\u304b\u3068\u3044\u3046\u3068\u3001\u3053\u306e\u53cd\u5fdc\u304cRuBP\u306e\u70ad\u7d20\u30fc\u70ad\u7d20\u7d50\u5408\u3092\u5207\u308b\u4ee3\u308f\u308a\u306b\u3001\u65b0\u3057\u3044\u70ad\u7d20\u30fc\u70ad\u7d20\u7d50\u5408\u3092\u751f\u3058\u308b\u53cd\u5fdc\u3060\u304b\u3089\u3067\u3059\u3002\u7b49\u4fa1\u4ea4\u63db\u3068\u3044\u3046\u308f\u3051\u3067\u3059\u3002 3-phosphoglycerate \u304b\u3089RuBP\u3092\u5408\u6210\u3059\u308b\u305f\u3081\u306b\u306f\u30a8\u30cd\u30eb\u30ae\u30fc\u304c\u5fc5\u8981\u306a\u306e\u3067\u3001\u5f53\u7136\u306a\u304c\u3089\u70ad\u9178\u56fa\u5b9a\u306f\u30a8\u30cd\u30eb\u30ae\u30fc\u304c\u306a\u3044\u3068\u9032\u307f\u307e\u305b\u3093\u3002<\/p>\n\n\n\n<h3>\u5149\u5408\u6210\u306e\u526f\u53cd\u5fdc\u3092\u9006\u306b\u5229\u7528\u3059\u308b<\/h3>\n\n\n\n<p>RubisCO\u306e\u53cd\u5fdc\u306f\u5730\u7403\u306e\u751f\u547d\u306e\u5fc3\u81d3\u90e8\u3068\u3082\u3044\u3048\u308b\u306e\u3067\u3059\u304c\u3001\u3053\u306e\u53cd\u5fdc\u306b\u306f\u5927\u304d\u306a\u554f\u984c\u304c\u3042\u308a\u307e\u3059\u3002\u305d\u308c\u306f\u3001\u5927\u6c17\u4e2d\u306b\u306f\u4e8c\u9178\u5316\u70ad\u7d20\u3068\u69cb\u9020\u304c\u985e\u4f3c\u3057\u3066\u3044\u308b\u9178\u7d20\u5206\u5b50\u304c\u5927\u91cf\u306b\u5b58\u5728\u3059\u308b\u3053\u3068\u3067\u3059\u3002\u5927\u6c17\u4e2d\u306e\u4e8c\u9178\u5316\u70ad\u7d20\u6fc3\u5ea6\u306f\u3001\u4e0a\u6607\u3057\u305f\u3068\u306f\u8a00\u3063\u3066\u3082400 ppm\u304f\u3089\u3044\u3057\u304b\u3042\u308a\u307e\u305b\u3093\u3002\u4e00\u65b9\u3001\u9178\u7d20\u306f20%\u3082\u5b58\u5728\u3057\u307e\u3059\u3002RubisCO\u306f\u3053\u306e\u5927\u91cf\u306b\u3042\u308b\u9178\u7d20\u3092\u907f\u3051\u306a\u304c\u3089\u4e8c\u9178\u5316\u70ad\u7d20\u3068\u53cd\u5fdc\u3059\u308b\u3068\u3044\u3046\u975e\u5e38\u306b\u9ad8\u5ea6\u306a\u3053\u3068\u3092\u3084\u3063\u3066\u3044\u308b\u306e\u3067\u3059\u3002\u3057\u304b\u3057\u3001\u4e00\u90e8\u306eRuBP\u306f\u9178\u7d20\u3068\u53cd\u5fdc\u3057\u3066\u3057\u307e\u3044\u307e\u3059\u3002\u3053\u306e\u5834\u5408\u306b\u751f\u6210\u3059\u308b\u306e\u304c\u30012-phosphoglycolate (2PG)\u3068\u3044\u3046\u7269\u8cea\u3067\u3001\u30b0\u30ea\u30b3\u30fc\u30eb\u9178\u304c\u30ea\u30f3\u9178\u5316\u3055\u308c\u305f\u3082\u306e\u3067\u3059\u3002\u591a\u304f\u306e\u5149\u5408\u6210\u306e\u7814\u7a76\u8005\u306f\u30012PG\u306e\u751f\u6210\u304c\u5149\u5408\u6210\u306e\u52b9\u7387\u3092\u4f4e\u4e0b\u3055\u305b\u308b\u3068\u3057\u3066\u3001\u3053\u308c\u3092\u767a\u751f\u3055\u305b\u306a\u3044\u305f\u3081\u306e\u7814\u7a76\u3092\u884c\u3063\u3066\u3044\u307e\u3059\u3002\u4e00\u65b9\u6211\u3005\u306f\u3001\u30b0\u30ea\u30b3\u30fc\u30eb\u9178\u304c\u30dd\u30ea\u30a8\u30b9\u30c6\u30eb\u3078\u3068\u5909\u63db\u53ef\u80fd\u306a\u69cb\u9020\u3092\u6301\u3063\u3066\u3044\u308b\u3053\u3068\u306b\u7740\u76ee\u3057\u307e\u3057\u305f\u3002RubisCO\u3092\u30dd\u30ea\u30de\u30fc\u5408\u6210\u306e\u305f\u3081\u306b\u4f7f\u3048\u306a\u3044\u304b\u3068\u8003\u3048\u305f\u306e\u3067\u3059\u3002<br>\u3000\u3053\u306e\u30a2\u30a4\u30c7\u30a3\u30a2\u3092\u691c\u8a3c\u3059\u308b\u305f\u3081\u3001RubisCO\u3092\u542b\u3080\u70ad\u9178\u56fa\u5b9a\u306e\u4ee3\u8b1d\u7d4c\u8def\uff08\u30ab\u30eb\u30d3\u30f3\u30b5\u30a4\u30af\u30eb\u306e\u4e00\u90e8\uff09\u3092\u6211\u3005\u304c\u30dd\u30ea\u30de\u30fc\u5408\u6210\u306b\u7528\u3044\u3066\u3044\u308b\u5927\u8178\u83cc\u306b\u5c0e\u5165\u3057\u307e\u3057\u305f\u3002\u3053\u308c\u3067\u3001\u5149\u5408\u6210\u751f\u7269\u306e\u4ee3\u8b1d\u306e\u4e00\u90e8\u3092\u6301\u3063\u3066\u3044\u308b\u5909\u306a\u5927\u8178\u83cc\u304c\u3067\u304d\u307e\u3057\u305f\u3002\u4e0a\u306b\u8aac\u660e\u3057\u305f\u3088\u3046\u306b\u3001\u30ab\u30eb\u30d3\u30f3\u30b5\u30a4\u30af\u30eb\u306f\u5149\u304c\u7121\u304f\u3066\u3082\u52d5\u304d\u307e\u3059\u3002\u5927\u8178\u83cc\u306e\u7d30\u80de\u5185\u3067\u3001\u30ab\u30eb\u30d3\u30f3\u30b5\u30a4\u30af\u30eb\u3092\u52d5\u304b\u3057\u3066\u3001\u30b0\u30ea\u30b3\u30fc\u30eb\u9178\u304c\u4f5c\u308c\u308b\u304b\u8a66\u3057\u307e\u3057\u305f\u3002\u305d\u306e\u7d50\u679c\u3001\u91cf\u306f\u975e\u5e38\u306b\u5c11\u306a\u3044\u3082\u306e\u306e\u3001\u30b0\u30ea\u30b3\u30fc\u30eb\u9178\u304c\u5408\u6210\u3055\u308c\u3001\u30dd\u30ea\u30de\u30fc\u5408\u6210\u306b\u3082\u6210\u529f\u3057\u307e\u3057\u305f\u3002\u73fe\u5728\u306f\u3001\u672c\u7d4c\u8def\u306e\u52b9\u7387\u6539\u5584\u306b\u53d6\u308a\u7d44\u3093\u3067\u3044\u307e\u3059\u3002<\/p>\n\n\n\n<figure class=\"wp-block-image\"><img decoding=\"async\" loading=\"lazy\" width=\"995\" height=\"699\" src=\"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-content\/uploads\/2020\/08\/RubisCOPathway.jpg\" alt=\"\" class=\"wp-image-545\" srcset=\"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-content\/uploads\/2020\/08\/RubisCOPathway.jpg 995w, https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-content\/uploads\/2020\/08\/RubisCOPathway-300x211.jpg 300w, https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-content\/uploads\/2020\/08\/RubisCOPathway-768x540.jpg 768w\" sizes=\"(max-width: 995px) 100vw, 995px\" \/><\/figure>\n","protected":false},"excerpt":{"rendered":"<p>\u5149\u5408\u6210\u3068\u306f \u5149\u5408\u6210\u306f\u3001\u5149\u306e\u30a8\u30cd\u30eb\u30ae\u30fc\u3092\u5229\u7528\u3057\u3066\u6c34\u3068\u4e8c\u9178\u5316\u70ad\u7d20\u304b\u3089\u6709\u6a5f\u7269\u3092\u5408\u6210\u3057\u307e\u3059\u3002\u8a00\u3046\u307e\u3067\u3082\u306a\u304f\u3001\u5730\u7403\u4e0a\u306e\u751f\u547d\u6d3b\u52d5\u3092\u7dad\u6301\u3059\u308b\u3046\u3048\u3067\u6700\u3082\u91cd\u8981\u306a\u53cd\u5fdc\u3067\u3059\u3002\u690d\u7269\u3067\u975e\u5e38\u306b\u767a\u9054\u3057\u3066\u3044\u307e\u3059\u304c\u3001\u7d30\u83cc\u306b\u3082\u5149\u5408\u6210\u80fd\u529b\u3092\u6301\u3064\u3082\u306e\u304c\u3044\u307e\u3059\u3002 &hellip;<br \/><a href=\"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/research\/topicrubisco.html\" class=\"more-link pen_button pen_element_default pen_icon_arrow_double\">\u7d9a\u304d\u3092\u8aad\u3080 <span class=\"screen-reader-text\">\u5149\u5408\u6210\u306e\u9375\u9175\u7d20RubisCO\u3092\u7528\u3044\u305f\u3082\u306e\u4f5c\u308a<\/span><\/a><\/p>\n","protected":false},"author":2,"featured_media":179,"comment_status":"closed","ping_status":"closed","sticky":false,"template":"","format":"standard","meta":[],"categories":[3],"tags":[],"_links":{"self":[{"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/posts\/148"}],"collection":[{"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/users\/2"}],"replies":[{"embeddable":true,"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/comments?post=148"}],"version-history":[{"count":9,"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/posts\/148\/revisions"}],"predecessor-version":[{"id":699,"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/posts\/148\/revisions\/699"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/media\/179"}],"wp:attachment":[{"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/media?parent=148"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/categories?post=148"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/c-mng.cwh.hokudai.ac.jp\/biosynchem.eng\/Root\/wp-json\/wp\/v2\/tags?post=148"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}